Maximum likelihood (ML) phylogenetic tree based on a concatenated dataset of ITS, cmdA, rpb2, and tub2. In total, 22 strains representing 19 taxa were included in the concatenated dataset, with 2424 characters (ITS 569 bp, cmdA 709 bp, rpb2 721 bp, and tub2 425 bp) including alignment gaps. The tree was rooted with Inaequalispora prestonii CBS 175.73. MLB ≥ 70% and BPPs ≥ 0.95 were shown at each node; values lower than these thresholds are indicated by a hyphen (–). The scale bar indicates the number of estimated substitutions per site. The strains introduced in this study are in red and novel species are in bold. The ex-type strains are marked with T.